Discovery of subspecies toxic to dipteran and coleopteran insects in 1977 and 1983, respectively.

1977年以前只发现13个，且这些都只对鳞翅目害有。

Any of various insect larvae that live and feed within a leaf, including caterpillars, sawfly larvae, beetle and weevil grubs, and dipteran maggots.

许多在叶中生活并取食昆幼称，包括毛、叶蜂幼、甲和象甲以及双翅类昆蛆。