2.Nutlets 4, obovoid, netted on back, areole nearly 1/2 as long as nutlet.
小坚4,倒卵球,在后面,小窠差不多1/2倍于小坚上得到。
3.Fruit dehiscent silicles or rarely siliques, obpyriform, obovoid, or rarely linear, shortly stipitate;
开裂短角少长角,倒梨,倒卵球,线少,具短柄;
4.Seeds obovoid-elliptic, base rostrate, apex retuse, raphe raised, ventral holes groovelike or obovate-elliptic in shape upward nearly 1/3.
种子倒卵球,基部具喙,先端微凹,种脊凸起,腹面洞无沟倒卵状达近上部1/3。
5.Tendril unbranched or bifurcate. Inflorescence usually a polychasium. Seeds elliptic, obovoid-elliptic, or obtriangular, surface smooth, corrugated, or with strumose protuberance or ribs.