subdominant

the subdominant male in a pride of lions.

狮群中仅次狮王的一头雄狮

Green bristlegrass was dominant species,and fat hen and cockspur grass were subdominant species.

研究区内有野植物25种,其中优势种群是狗尾种群,亚优势种群为灰菜、稗种群。

The plagal cadence, sometimes known as the Amen cadence from ecclesiastical use, is the progression from the chord of the subdominant to the tonic, IV - I.

变格终止，有时也被称为源教会的“阿们终止”，是指从下属音和到主音和的进行，IV–I。