2.Leaves simple, alternate or opposite, exstipulate, margin entire or dentate, venation pinnate or triplinerved.
单叶,互生或对生,无托叶,边缘全缘或具牙齿,羽状脉或离基三出脉。
3.Leaf blade narrowly elliptic-lanceolate to ovate-lanceolate, margin spiniform dentate; bracts of cupule subulate, ligulate, or linear, often reflexed.